LIGHT AND DARK ADAPTATION Physiology LectureLIGHT AND DARK ADAPTATION Physiology Lecture

RETINA

 Light sensitive portion of the eye
 Contains cones for color vision
 Contains rods for night vision
 Contains neural architecture
 Light must pass through the neural elements to
strike
 The light sensitive rods and cones

LIGHT ADAPTATION

 Bright light for some time (hrs)
MECHANISMS
 Photosensitive chemicals in rods and cones will be
reduced to retinals and opsins
 Retinals to Vitamin A
 concentration of photosensitive chemicals reduced

  • Change from rod to cone
    vision
  • Constriction of pupil
     Less light enters
  • Neural adaptation
     Diminution of signals

DARK ADAPTATION

 Darkness for a long time
 Retinals and opsins back to light sensitive pigment
 Vitamin A back to retinal
 Increase in light-sensitive pigments

When person is exposed to total darkness
 After remaining in light for several hours
 Sensitivity of the retina—- increased
 Process by which one can see in dim light

 Maximum adaptation time
 very low— on entering the darkness
 1 min—-10 fold
 20 min—–6000 fold
 40 min—-25,000 fold

Dark adaptation curve
 Adaptation of the cones
 Early portion of the curve
 Adaptation occurs four times more rapidly in
cones than in rods
 Less level of sensitivity—Cease adapting after a few
minutes

Dark adaptation curve
 Adaptation by rods
 slow adaptation
 More sensitive—Continue to adapt for
minutes/hours
 Rods summate to increase their sensitivity
 Convergence of 100 or more rods onto a single
ganglion cells in the retina

Other Mechanisms of Adaptation
 Change in pupillary size
 Pupil dilate more light enters
 Adaptation of 30 fold— sec
 Neural adaptation
 Enhancement of signals
 Degree of adaptation is few fold

Importance of Dark and Light Adaptation

 Adaptation—-500,000 to 1 million times
 The detection of images on the retina is a function
of discriminating between dark and light spots.
 Registering of images by retina
 Detection of dark and light spots
 Receptors respond to lighter areas but not to darker
areas

 Enter the sun from a movie theater, even the dark
spots appear bright leaving little contrast
 Entire visual image is bleached
 Enter darkness from light, the sensitivity of retina is
so slight—light spots are not light enough to
register (cannot excite the retina)
 After dark adaptation the light spots begin to
register

 Eyes can function in
 Bright sunlight— light adaptation
 Star light—dark adaptation

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